| 1974 - 1980 | Medical studies, University of Bern. | 
| 1981 - 1982 | Research Associate at the Institute for Biomedical Engineering, Federal Institute of Technology (ETH) and University of Zurich (Prof. Dr. F. H. Walz). | 
| 1983 | Resident at the Department of Internal Medicine, Hospital of Sumiswald / Bern (Dr. M. Fey).
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| 1984 | Research Associate at the Institute for Biomedical Engineering, Federal Institute of Technology (ETH) and University of Zurich (Prof. Dr. F. H. Walz). | 
| 1985 - 1988 | Research Associate at the Department of Physiology, University of Bern (Profs. S. Weidmann and R. Weingart). | 
| 1988 - 1989 | Research Assistant Professor at the Department of Physiology, University of Maryland UMAB (Prof. Dr. W. J. Lederer). | 
| 1990 | Research Assistant Professor at the Department of Physiology, Medical Biotechnology Center UMBI (Prof. Dr. W. J. Lederer). | 
| 1990 - 1995 | Assistant Professor (Oberassistent) at the Department of Physiology, University of Bern. Cellular regulatory mechanisms in cardiac contraction and relaxation: role of the Na/Ca exchanger. | 
| 1996 - 2000 | Associate Professor at the Department of Physiology, University of Bern. | 
| 2000 | Visiting Professor, Department of Physiology, Loyola University, Maywood IL, USA. | 
| 2000 - today | Professor at the Department of Physiology, University of Bern. | 
| 2007 | Visiting Professor, Department of Pharmacology and Physiology, UMDNJ, Newark, NJ, USA. | 
| 2013 | Visiting Professor, Department of Pharmacology and Physiology, Rutgers University, Newark, NJ, USA. | 
| 1990 - 1994 | Lectures for pharmacy students: cardiovascular physiology | 
| 1990 - today | Practical courses for medical students | 
| 1995 - 1997 | Lectures for medical students: renal Physiology, energy metabolism and temperature regulation | 
| 1997 - today | Lectures for students of veterinary medicine and pharmacy: cardiovascular physiology | 
| 1999 - 2008 | Coordinator, lecturer and tutor of the renal PBL course for medical students and dentists. Tutor of the PBL course "Metabolism". | 
| 2009 - today | Coordinator, lecturer and tutor of the cardiovascular and respiration PBL course for medical students and dentists. | 
| 2009 - today | Coordinator and lecturer of the module “Heart”, Master
course in Biomedical Sciences (BMSc) |