| 1975 - 1981 | Medical studies at the University of Bern |
| 1980 - 1981 | Internship at the District Hospital Litembo, Tanzania |
| 1984 | M. D. at the Faculty of Medicine in Bern ( Thesis with Prof. Weidmann) |
| 1985 - 1987 | Postdoc at the Max Planck Institute for Psychiatry in Munich (Prof. Lux) |
| 1987 - 1989 | Postdoc at the Institute of Physiology in Bern (Prof. Lüscher) |
| 1990 - | "Dozent" at the Institute of Physiology in Bern |
| 1994 | Habilitation at the Faculty of Medicine in Bern |
| 2002 | Associate Professor at the Institute of Physiology in Bern |
